Justin Long

Alvin Seville (Voice)

Matthew Gray Gubler

Simon Seville (Voice)

Jesse McCartney

Theodore Seville (Voice)

Christina Applegate

Brittany Miller (Voice)

Anna Faris

Jeanette Miller (Voice)

Vanessa Bagdasarian

Eleanor Miller / Lucy/ Kate (Voice)

Ross Bagdasarian, Jr.

Dave Seville (Voice)